About the Authors

Godfrey Kigozi

gkigozi@rhsp.org

Affiliation Rakai Health Sciences Program, Entebbe, Uganda

Richard Musoke

Affiliation Rakai Health Sciences Program, Entebbe, Uganda

Stephen Watya

Affiliation Rakai Health Sciences Program, Entebbe, Uganda

Nehemia Kighoma

Affiliation Rakai Health Sciences Program, Entebbe, Uganda

James Nkale

Affiliation Rakai Health Sciences Program, Entebbe, Uganda

Mary Nakafeero

Affiliation Rakai Health Sciences Program, Entebbe, Uganda

Dan Namuguzi

Affiliation Department of Urology, Mulago Hospital, Kampala, Uganda

David Serwada

Affiliations Rakai Health Sciences Program, Entebbe, Uganda, School of Public Health, Makerere University College of Health Sciences, Kampala, Uganda

Fred Nalugoda

Affiliation Rakai Health Sciences Program, Entebbe, Uganda

Nelson Sewankambo

Affiliations Rakai Health Sciences Program, Entebbe, Uganda, College of Health Sciences, Makerere University, Kampala, Uganda

Maria Joan Wawer

Affiliations Rakai Health Sciences Program, Entebbe, Uganda, Department of Epidemiology, Johns Hopkins University, Bloomberg School of Public Health, Baltimore, Maryland, United States of America

Ronald Henry Gray

Affiliations Rakai Health Sciences Program, Entebbe, Uganda, Department of Epidemiology, Johns Hopkins University, Bloomberg School of Public Health, Baltimore, Maryland, United States of America

Competing Interests

Ronald H. Gray is a member of the PLOS ONE editorial board. This does not alter the authors' adherence to PLOS ONE Editorial policies and criteria.

Author Contributions

Conceived and designed the experiments: RHG GK MJW RM SW DS NS. Performed the experiments: GK NK JN SW DN MN FN. Analyzed the data: RM GK MN RHG JN. Wrote the paper: GK RHG RM MJW FN DS NS SW DN NK JN MN.