About the Authors

Jenni Liljavirta

Affiliation Department of Veterinary Biosciences, University of Helsinki, Helsinki, Finland

Mikael Niku

Affiliation Department of Veterinary Biosciences, University of Helsinki, Helsinki, Finland

Tiina Pessa-Morikawa

Affiliation Department of Veterinary Biosciences, University of Helsinki, Helsinki, Finland

Anna Ekman

Affiliation Department of Veterinary Biosciences, University of Helsinki, Helsinki, Finland

Antti Iivanainen

antti.iivanainen@helsinki.fi

Affiliation Department of Veterinary Biosciences, University of Helsinki, Helsinki, Finland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AI JL MN AE. Performed the experiments: JL MN AE TPM. Analyzed the data: AI JL MN AE. Contributed reagents/materials/analysis tools: AI JL MN. Wrote the paper: JL MN AI TPM AE.