About the Authors

Sally D. Hofmeyr

salhofmeyr@gmail.com

Affiliations School of Animal, Plant and Environmental Sciences, University of the Witwatersrand, Johannesburg, Gauteng, South Africa, Animal Demography Unit, Department of Biological Sciences, University of Cape Town, Cape Town, Western Cape, South Africa

Craig T. Symes

Affiliation School of Animal, Plant and Environmental Sciences, University of the Witwatersrand, Johannesburg, Gauteng, South Africa

Leslie G. Underhill

Affiliation Animal Demography Unit, Department of Biological Sciences, University of Cape Town, Cape Town, Western Cape, South Africa

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SDH LGU. Performed the experiments: SDH. Analyzed the data: SDH. Contributed reagents/materials/analysis tools: LGU. Wrote the paper: SDH CTS LGU. Designed the statistical model used in this analysis: LGU.