About the Authors

Claire W. Varian-Ramos

claire.ramos@colostate-pueblo.edu

Affiliation Biology Department, Colorado State University – Pueblo, Pueblo, Colorado, United States of America

John P. Swaddle

Affiliation Institute for Integrative Bird Behavior Studies, Biology Department, The College of William and Mary, Williamsburg, Virginia, United States of America

Daniel A. Cristol

Affiliation Institute for Integrative Bird Behavior Studies, Biology Department, The College of William and Mary, Williamsburg, Virginia, United States of America

Competing Interests

The authors acknowledge that this study was funded in part by E.I. DuPont de Nemours and Company (DuPont). Neither DuPont nor any employee of DuPont was involved in the experimental design, data collection, analysis and interpretation, or preparation of this manuscript.

Author Contributions

Conceived and designed the experiments: CWVR JPS DAC. Performed the experiments: CWVR. Analyzed the data: CWVR. Contributed reagents/materials/analysis tools: JPS DAC. Wrote the paper: CWVR JPS DAC.