About the Authors

William C. Bowen

Affiliation Department of Pathology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Amantha W. Michalopoulos

Affiliation Department of Pathology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Anne Orr

Affiliation Department of Pathology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Michael Q. Ding

Affiliation Department of Pathology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Donna B. Stolz

Affiliation Department of Cell Biology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

George K. Michalopoulos

michalopoulosgk@msx.upmc.edu

Affiliation Department of Pathology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GKM WCB. Performed the experiments: AWM AO MQD DBS. Analyzed the data: GKM. Contributed reagents/materials/analysis tools: DBS. Wrote the paper: GKM DBS.