About the Authors

Lei Fang

Contributed equally to this work with: Lei Fang, Ruiping Tian

Affiliation National Key Laboratory of Crop Genetics and Germplasm Enhancement, Cotton Hybrid R & D Engineering Center (the Ministry of Education), Nanjing Agricultural University, Nanjing, China

Ruiping Tian

Contributed equally to this work with: Lei Fang, Ruiping Tian

Affiliation National Key Laboratory of Crop Genetics and Germplasm Enhancement, Cotton Hybrid R & D Engineering Center (the Ministry of Education), Nanjing Agricultural University, Nanjing, China

Jiedan Chen

Affiliation National Key Laboratory of Crop Genetics and Germplasm Enhancement, Cotton Hybrid R & D Engineering Center (the Ministry of Education), Nanjing Agricultural University, Nanjing, China

Sen Wang

Affiliation National Key Laboratory of Crop Genetics and Germplasm Enhancement, Cotton Hybrid R & D Engineering Center (the Ministry of Education), Nanjing Agricultural University, Nanjing, China

Xinghe Li

Affiliation National Key Laboratory of Crop Genetics and Germplasm Enhancement, Cotton Hybrid R & D Engineering Center (the Ministry of Education), Nanjing Agricultural University, Nanjing, China

Peng Wang

Affiliation National Key Laboratory of Crop Genetics and Germplasm Enhancement, Cotton Hybrid R & D Engineering Center (the Ministry of Education), Nanjing Agricultural University, Nanjing, China

Tianzhen Zhang

cotton@njau.edu.cn

Affiliation National Key Laboratory of Crop Genetics and Germplasm Enhancement, Cotton Hybrid R & D Engineering Center (the Ministry of Education), Nanjing Agricultural University, Nanjing, China

Competing Interests

Tianzhen Zhang is a PLOS ONE Editorial Board member, this does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: TZ. Performed the experiments: LF RT SW XL PW. Analyzed the data: LF JC. Wrote the paper: LF TZ.