About the Authors

Marie T. Dittmann

marie.dittmann@agrl.ethz.ch

Affiliations Clinic for Zoo Animals, Exotic Pets and Wildlife, Vetsuisse Faculty Zurich, University of Zurich, Zurich, Switzerland, ETH Zurich, Institute for Agricultural Sciences, Zurich, Switzerland

Ullrich Runge

Affiliation Kamelhof Olmerswil, Neukirch/Thur, Switzerland

Richard A. Lang

Affiliation Cochranton Veterinary Hospital, Cochranton, Pennsylvania, United States of America

Dario Moser

Affiliation Zoological Institute, University of Basel, Basel, Switzerland

Cordula Galeffi

Affiliation Zurich Zoo, Zurich, Switzerland

Michael Kreuzer

Affiliation ETH Zurich, Institute for Agricultural Sciences, Zurich, Switzerland

Marcus Clauss

Affiliation Clinic for Zoo Animals, Exotic Pets and Wildlife, Vetsuisse Faculty Zurich, University of Zurich, Zurich, Switzerland

Competing Interests

Co-author Ullrich Runge is employed by Kamelhof Olmerswil, a small private farm keeping camels. This does not alter our adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MC MK. Performed the experiments: MTD UR RAL DM CG MC. Analyzed the data: MTD MC. Wrote the paper: MTD MC. Manuscript review: UR RAL DM CG MK. Animal husbandry: UR RAL DM.