About the Authors

Shannon E. Boye

jacobsos@mail.med.upenn.edu (SGJ); shaire@ufl.edu (SEB)

Affiliation Department of Ophthalmology, College of Medicine, University of Florida, Gainesville, Florida, United States of America

Wei-Chieh Huang

Affiliation Scheie Eye Institute, Department of Ophthalmology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Alejandro J. Roman

Affiliation Scheie Eye Institute, Department of Ophthalmology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Alexander Sumaroka

Affiliation Scheie Eye Institute, Department of Ophthalmology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Sanford L. Boye

Affiliation Department of Ophthalmology, College of Medicine, University of Florida, Gainesville, Florida, United States of America

Renee C. Ryals

Affiliation Department of Ophthalmology, College of Medicine, University of Florida, Gainesville, Florida, United States of America

Melani B. Olivares

Affiliation Scheie Eye Institute, Department of Ophthalmology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Qing Ruan

Affiliation Department of Ophthalmology, College of Medicine, University of Florida, Gainesville, Florida, United States of America

Budd A. Tucker

Affiliation Stephen A. Wynn Institute for Vision Research, University of Iowa Carver College of Medicine, Iowa City, Iowa, United States of America

Edwin M. Stone

Affiliations Stephen A. Wynn Institute for Vision Research, University of Iowa Carver College of Medicine, Iowa City, Iowa, United States of America, Howard Hughes Medical Institute, University of Iowa Carver College of Medicine, Iowa City, Iowa, United States of America

Anand Swaroop

Affiliation Neurobiology-Neurodegeneration & Repair Laboratory, National Eye Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Artur V. Cideciyan

Affiliation Scheie Eye Institute, Department of Ophthalmology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

William W. Hauswirth

Affiliation Department of Ophthalmology, College of Medicine, University of Florida, Gainesville, Florida, United States of America

Samuel G. Jacobson

jacobsos@mail.med.upenn.edu (SGJ); shaire@ufl.edu (SEB)

Affiliation Scheie Eye Institute, Department of Ophthalmology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SGJ SEB AVC A. Swaroop EMS WWH. Performed the experiments: WCH AJR A. Sumaroka SLB RCR MBO QR BAT. Analyzed the data: WCH AJR A. Sumaroka SLB RCR MBO QR BAT. Wrote the paper: SGJ SEB AVC.