About the Authors

Fumiyoshi Yamazaki

Affiliation Section on Neuroendocrinology, Program in Developmental Endocrinology and Genetics, The Eunice Shriver Kennedy National Institute of Child Health and Human Development, National Institutes of Health, Bethesda, Maryland, United States of America

Hyun Hee Kim

Affiliation Section on Neuroendocrinology, Program in Developmental Endocrinology and Genetics, The Eunice Shriver Kennedy National Institute of Child Health and Human Development, National Institutes of Health, Bethesda, Maryland, United States of America

Pierre Lau

Current address: Laboratory for the Research of Neurodegenerative Diseases, Department of Molecular and Developmental Genetics, University of Leuven, Flemish Brabant, Belgium

Affiliation Division of intramural research, National Institute of Neurological Disorders and Stroke, National Institutes of Health, Bethesda, Maryland, United States of America

Christopher K. Hwang

Affiliation Departments of Ophthalmology and Pharmacology, Emory University School of Medicine, Atlanta, Georgia, United States of America

P. Michael Iuvone

Affiliation Departments of Ophthalmology and Pharmacology, Emory University School of Medicine, Atlanta, Georgia, United States of America

David Klein

kleind@mail.nih.gov

Affiliation Section on Neuroendocrinology, Program in Developmental Endocrinology and Genetics, The Eunice Shriver Kennedy National Institute of Child Health and Human Development, National Institutes of Health, Bethesda, Maryland, United States of America

Samuel J. H. Clokie

Current address: West Midlands Regional Genetics Laboratories, Birmingham Women's NHS (National Health Service) Foundation Trust, Birmingham, West Midlands, United Kingdom

Affiliation Section on Neuroendocrinology, Program in Developmental Endocrinology and Genetics, The Eunice Shriver Kennedy National Institute of Child Health and Human Development, National Institutes of Health, Bethesda, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SJHC DCK PMI. Performed the experiments: SJHC HHK PL CKH FY. Analyzed the data: FY HHK PL CKH PMI DCK SJHC. Contributed reagents/materials/analysis tools: FY HHK PL CKH SJHC. Wrote the paper: SJHC DCK PMI FY.