About the Authors

Jing Huang

Contributed equally to this work with: Jing Huang, Xiangming Li, Jordana G. A. Coelho-dos-Reis

Affiliation HIV and Malaria Vaccine Program, Aaron Diamond AIDS Research Center, Affiliate of The Rockefeller University, New York, New York, United States of America

Xiangming Li

Contributed equally to this work with: Jing Huang, Xiangming Li, Jordana G. A. Coelho-dos-Reis

Affiliation HIV and Malaria Vaccine Program, Aaron Diamond AIDS Research Center, Affiliate of The Rockefeller University, New York, New York, United States of America

Jordana G. A. Coelho-dos-Reis

Contributed equally to this work with: Jing Huang, Xiangming Li, Jordana G. A. Coelho-dos-Reis

Affiliation HIV and Malaria Vaccine Program, Aaron Diamond AIDS Research Center, Affiliate of The Rockefeller University, New York, New York, United States of America

James M. Wilson

Affiliation Gene Therapy Program, Department of Pathology and Laboratory Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Moriya Tsuji

mtsuji@adarc.org

Affiliation HIV and Malaria Vaccine Program, Aaron Diamond AIDS Research Center, Affiliate of The Rockefeller University, New York, New York, United States of America

Competing Interests

This work was partly funded by Otsuka Pharmaceutical Co. Ltd. James M. Wilson is a consultant to ReGenX Holdings, and is a founder of, holds equity in, and receives a grant from affiliates of ReGenX Holdings; in addition, he is an inventor on patents licensed to various biopharmaceutical companies, including affiliates of ReGenX Holdings. There are no further patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors. Furthermore, there are no declared patents that relate directly to this study.

Author Contributions

Conceived and designed the experiments: MT XL JH JGCR. Performed the experiments: XL JH JGCR. Analyzed the data: MT XL JH JGCR. Contributed reagents/materials/analysis tools: JMW. Wrote the paper: MT XL JH JMW JGCR.