About the Authors

Ryohei Numata

Affiliation Department of Biomedical Engineering, Faculty of Life and Medical Sciences, Doshisha University, Kyotanabe, Japan

Naoki Okumura

Affiliations Department of Biomedical Engineering, Faculty of Life and Medical Sciences, Doshisha University, Kyotanabe, Japan, Department of Ophthalmology, Kyoto Prefectural University of Medicine, Kyoto, Japan

Makiko Nakahara

Affiliation Department of Biomedical Engineering, Faculty of Life and Medical Sciences, Doshisha University, Kyotanabe, Japan

Morio Ueno

Affiliation Department of Ophthalmology, Kyoto Prefectural University of Medicine, Kyoto, Japan

Shigeru Kinoshita

Affiliation Department of Ophthalmology, Kyoto Prefectural University of Medicine, Kyoto, Japan

Daisuke Kanematsu

Affiliation Division of Regenerative Medicine, Institute for Clinical Research, Osaka National Hospital, National Hospital Organization, Osaka, Japan

Yonehiro Kanemura

Affiliation Division of Regenerative Medicine, Institute for Clinical Research, Osaka National Hospital, National Hospital Organization, Osaka, Japan

Yoshiki Sasai

Affiliation RIKEN Center for Developmental Biology, Hyogo, Japan

Noriko Koizumi

nkoizumi@mail.doshisha.ac.jp

Affiliation Department of Biomedical Engineering, Faculty of Life and Medical Sciences, Doshisha University, Kyotanabe, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NO MU SK YS NK. Performed the experiments: RN MN. Analyzed the data: RN MN NO NK. Contributed reagents/materials/analysis tools: SK DK YK YS NK. Wrote the paper: NO NK.