About the Authors

Takanori Nihira

Affiliation Graduate School of Science and Technology, Niigata University, Niigata, Japan

Yuka Saito

Affiliation Graduate School of Science and Technology, Niigata University, Niigata, Japan

Ken’ichi Ohtsubo

Affiliation Graduate School of Science and Technology, Niigata University, Niigata, Japan

Hiroyuki Nakai

mkitaoka@affrc.go.jp (MK); nakai@agr.niigata-u.ac.jp (HN)

Affiliation Graduate School of Science and Technology, Niigata University, Niigata, Japan

Motomitsu Kitaoka

mkitaoka@affrc.go.jp (MK); nakai@agr.niigata-u.ac.jp (HN)

Affiliation National Food Research Institute, National Agriculture and Food Research Organization, Tsukuba, Ibaraki, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HN MK. Performed the experiments: TN YS MK. Analyzed the data: TN YS HN MK. Contributed reagents/materials/analysis tools: KO HN MK. Wrote the paper: TN HN MK.