About the Authors

Thomas J. O’Shea

osheat@usgs.gov

Affiliation United States Geological Survey, Fort Collins Science Center, Fort Collins, Colorado, United States of America

Richard A. Bowen

Affiliation Department of Biomedical Sciences, Colorado State University, Fort Collins, Colorado, United States of America

Thomas R. Stanley

Affiliation United States Geological Survey, Fort Collins Science Center, Fort Collins, Colorado, United States of America

Vidya Shankar

Current address: U.S. Army Public Health Command, Joint Base Lewis-McChord, Fort Lewis, Washington, United States of America

Affiliations Department of Biomedical Sciences, Colorado State University, Fort Collins, Colorado, United States of America, Centers for Disease Control and Prevention, Atlanta, Georgia, United States of America

Charles E. Rupprecht

Affiliations Ross University School of Veterinary Medicine, Basseterre, Saint Kitts, West Indies, The Global Alliance for Rabies Control, Manhattan, Kansas, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CER RAB TJO VS. Performed the experiments: VS CER RAB TJO. Analyzed the data: TRS TJO VS. Contributed reagents/materials/analysis tools: CER RAB TJO VS. Wrote the paper: TJO RAB TRS VS CER.