About the Authors

Alyssa N. Malone

Affiliation Department of Microbiology, Immunology, and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Darcy M. Fletcher

Affiliation Department of Microbiology, Immunology, and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Megan B. Vogt

Affiliation Department of Microbiology, Immunology, and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Stephen K. Meyer

Affiliation Department of Microbiology, Immunology, and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Ann M. Hess

Affiliation Department of Statistics, Colorado State University, Fort Collins, Colorado, United States of America

Torsten M. Eckstein

eckstein@mail.colostate.edu

Affiliation Department of Microbiology, Immunology, and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TME. Performed the experiments: ANM DMF MBV SKM. Analyzed the data: ANM AMH TME. Wrote the manuscript: TME.