About the Authors

Yan Xu

Affiliations School of Life Sciences and Technology, Tongji University, Shanghai, People’s Republic of China, Department of Genome Biology, John Curtin School of Medical Research, The Australian National University, Canberra, Australia, State Key Laboratory of Pharmaceutical Biotechnology, Nanjing University, Nanjing, People’s Republic of China

Stephen J. Ohms

Affiliation Australian Cancer Research Foundation Biomolecular Resource Facility, John Curtin School of Medical Research, The Australian National University, Canberra, Australia

Zhen Li

Affiliation School of Life Sciences and Technology, Tongji University, Shanghai, People’s Republic of China

Qiao Wang

Affiliation National Information and Communications Technology Australia, Victoria Research Laboratory, The University of Melbourne, Melbourne, Australia

Guangming Gong

Affiliation State Key Laboratory of Pharmaceutical Biotechnology, Nanjing University, Nanjing, People’s Republic of China

Yiqiao Hu

Affiliation State Key Laboratory of Pharmaceutical Biotechnology, Nanjing University, Nanjing, People’s Republic of China

Zhiyong Mao

jun.fan@anu.edu.au (JYF); Frances.Shannon@canberra.edu.au (MFS); zhiyong_mao@tongji.edu.cn (ZM)

Affiliation School of Life Sciences and Technology, Tongji University, Shanghai, People’s Republic of China

M. Frances Shannon

jun.fan@anu.edu.au (JYF); Frances.Shannon@canberra.edu.au (MFS); zhiyong_mao@tongji.edu.cn (ZM)

Affiliations Department of Genome Biology, John Curtin School of Medical Research, The Australian National University, Canberra, Australia, University of Canberra, Canberra, Australia

Jun Y. Fan

jun.fan@anu.edu.au (JYF); Frances.Shannon@canberra.edu.au (MFS); zhiyong_mao@tongji.edu.cn (ZM)

Affiliation Department of Genome Biology, John Curtin School of Medical Research, The Australian National University, Canberra, Australia

Competing Interests

The authors would like to declare that we received funding from a commercial source, i.e. Bioplatforms Australia. This does not alter the authors' adherence to all PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: YX JYF MFS YH. Performed the experiments: YX ZL GG. Analyzed the data: SJO YX QW. Wrote the manuscript: JYF YX SJO. Data discussion, manuscript revision: MFS ZM.