About the Authors

Elisabeth Bowers

Affiliations Mucosal and Vaccine Research Program Colorado (MAVRC), University of Colorado Denver, Aurora, Colorado, United States of America, Infectious Disease Division, University of Colorado Denver, Aurora, Colorado, United States of America, Department of Microbiology, University of Colorado Denver, Aurora, Colorado, United States of America, Denver Veterans Affairs Medical Center, Denver, Colorado, United States of America

Ronald W. Scamurra

Affiliation Minneapolis VA Health Care System, University of Minnesota, Minneapolis, Minnesota, United States of America

Anil Asrani

Affiliation Minneapolis VA Health Care System, University of Minnesota, Minneapolis, Minnesota, United States of America

Lydie Beniguel

Affiliation GIMAP EA 3064, Faculté de Médecine, Université Jean Monnet, Saint Etienne, France

Samantha MaWhinney

Affiliations Mucosal and Vaccine Research Program Colorado (MAVRC), University of Colorado Denver, Aurora, Colorado, United States of America, Colorado School of Public Health, University of Colorado Denver, Aurora, Colorado, United States of America

Kathryne M. Keays

Affiliations Mucosal and Vaccine Research Program Colorado (MAVRC), University of Colorado Denver, Aurora, Colorado, United States of America, Infectious Disease Division, University of Colorado Denver, Aurora, Colorado, United States of America

Joseph R. Thurn

Affiliation Minneapolis VA Health Care System, University of Minnesota, Minneapolis, Minnesota, United States of America

Edward N. Janoff

* , e-mail:Edward.Janoff@ucdenver.edu

Affiliations Mucosal and Vaccine Research Program Colorado (MAVRC), University of Colorado Denver, Aurora, Colorado, United States of America, Infectious Disease Division, University of Colorado Denver, Aurora, Colorado, United States of America, Department of Microbiology, University of Colorado Denver, Aurora, Colorado, United States of America, Denver Veterans Affairs Medical Center, Denver, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EB RWS LB ENJ. Performed the experiments: EB RWS AA KMK. Analyzed the data: EB RWS SM. Contributed reagents/materials/analysis tools: JRT. Wrote the paper: EB ENJ. Patient recruitment: JRT.