About the Authors
- Lily Paemka
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Affiliations The University of Iowa, Iowa City, Iowa, United States of America, Department of Pediatrics, The University of Iowa, Iowa City, Iowa, United States of America, Interdisciplinary Program in Genetics, The University of Iowa, Iowa City, Iowa, United States of America
- Vinit B. Mahajan
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Affiliations The University of Iowa, Iowa City, Iowa, United States of America, Department of Ophthalmology and Visual Sciences, The University of Iowa, Iowa City, Iowa, United States of America, Roy J. and Lucille A. Carver College of Medicine, The University of Iowa, Iowa City, Iowa, United States of America
- Jessica M. Skeie
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Affiliations The University of Iowa, Iowa City, Iowa, United States of America, Department of Ophthalmology and Visual Sciences, The University of Iowa, Iowa City, Iowa, United States of America
- Levi P. Sowers
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Affiliations The University of Iowa, Iowa City, Iowa, United States of America, Department of Neurology, The University of Iowa, Iowa City, Iowa, United States of America, Roy J. and Lucille A. Carver College of Medicine, The University of Iowa, Iowa City, Iowa, United States of America
- Salleh N. Ehaideb
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Affiliations The University of Iowa, Iowa City, Iowa, United States of America, Interdisciplinary Program in Genetics, The University of Iowa, Iowa City, Iowa, United States of America, Department of Biology, The University of Iowa, Iowa City, Iowa, United States of America
- Pedro Gonzalez-Alegre
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Affiliations The University of Iowa, Iowa City, Iowa, United States of America, Department of Neurology, The University of Iowa, Iowa City, Iowa, United States of America, Roy J. and Lucille A. Carver College of Medicine, The University of Iowa, Iowa City, Iowa, United States of America, Interdisciplinary Graduate Program of Neuroscience, The University of Iowa, Iowa City, Iowa, United States of America
- Toshikuni Sasaoka
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Affiliation Center for Bioresources, Brain Research Institute, Niigata University, Niigata, Japan
- Hirotaka Tao
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Affiliation Hospital for Sick Kids, University of Toronto, Toronto, Canada
- Asuka Miyagi
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Affiliation Developmental Biology Department, National Institute for Basic Biology, Okazaki City, Japan
- Naoto Ueno
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Affiliation Developmental Biology Department, National Institute for Basic Biology, Okazaki City, Japan
- Keizo Takao
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Affiliations Section of Behavior Patterns, Center for Genetic Analysis of Behavior National Institute for Physiological Sciences, Okazaki, Japan, Japan Science and Technology Agency, Kawaguchi-shi, Japan
- Tsuyoshi Miyakawa
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Affiliations Section of Behavior Patterns, Center for Genetic Analysis of Behavior National Institute for Physiological Sciences, Okazaki, Japan, Japan Science and Technology Agency, Kawaguchi-shi, Japan, Division of Systems Medical Science, Institute for Comprehensive Medical Science, Fujita Health University, Toyoake, Japan
- Shu Wu
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Affiliations The University of Iowa, Iowa City, Iowa, United States of America, Department of Pediatrics, The University of Iowa, Iowa City, Iowa, United States of America
- Benjamin W. Darbro
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Affiliations The University of Iowa, Iowa City, Iowa, United States of America, Roy J. and Lucille A. Carver College of Medicine, The University of Iowa, Iowa City, Iowa, United States of America, Department of Pathology, The University of Iowa, Iowa City, Iowa, United States of America
- Polly J. Ferguson
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Affiliations The University of Iowa, Iowa City, Iowa, United States of America, Department of Pediatrics, The University of Iowa, Iowa City, Iowa, United States of America, Roy J. and Lucille A. Carver College of Medicine, The University of Iowa, Iowa City, Iowa, United States of America
- Andrew A. Pieper
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Affiliations The University of Iowa, Iowa City, Iowa, United States of America, Department of Neurology, The University of Iowa, Iowa City, Iowa, United States of America, Roy J. and Lucille A. Carver College of Medicine, The University of Iowa, Iowa City, Iowa, United States of America, Department of Psychiatry, The University of Iowa, Iowa City, Iowa, United States of America
- Jeremiah K. Britt
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Affiliations The University of Iowa, Iowa City, Iowa, United States of America, Roy J. and Lucille A. Carver College of Medicine, The University of Iowa, Iowa City, Iowa, United States of America, Department of Psychiatry, The University of Iowa, Iowa City, Iowa, United States of America
- John A. Wemmie
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Affiliations The University of Iowa, Iowa City, Iowa, United States of America, Interdisciplinary Graduate Program of Neuroscience, The University of Iowa, Iowa City, Iowa, United States of America, Department of Psychiatry, The University of Iowa, Iowa City, Iowa, United States of America
- Danielle S. Rudd
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Affiliations The University of Iowa, Iowa City, Iowa, United States of America, Interdisciplinary Program in Genetics, The University of Iowa, Iowa City, Iowa, United States of America, Department of Psychiatry, The University of Iowa, Iowa City, Iowa, United States of America
- Thomas Wassink
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Affiliations The University of Iowa, Iowa City, Iowa, United States of America, Roy J. and Lucille A. Carver College of Medicine, The University of Iowa, Iowa City, Iowa, United States of America, Department of Psychiatry, The University of Iowa, Iowa City, Iowa, United States of America
- Hatem El-Shanti
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Affiliations The University of Iowa, Iowa City, Iowa, United States of America, Department of Pediatrics, The University of Iowa, Iowa City, Iowa, United States of America, Roy J. and Lucille A. Carver College of Medicine, The University of Iowa, Iowa City, Iowa, United States of America, Shafallah Medical Genetics Center, Doha, Qatar
- Heather C. Mefford
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Affiliation Division of Genetic Medicine, Department of Pediatrics, University of Washington, Seattle, Washington, United States of America
- Gemma L. Carvill
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Affiliation Division of Genetic Medicine, Department of Pediatrics, University of Washington, Seattle, Washington, United States of America
- J. Robert Manak
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Affiliations The University of Iowa, Iowa City, Iowa, United States of America, Department of Pediatrics, The University of Iowa, Iowa City, Iowa, United States of America, Interdisciplinary Program in Genetics, The University of Iowa, Iowa City, Iowa, United States of America, Department of Biology, The University of Iowa, Iowa City, Iowa, United States of America, Interdisciplinary Graduate Program in Molecular and Cellular Biology, The University of Iowa, Iowa City, Iowa, United States of America
- Alexander G. Bassuk
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* E-mail: alexander-bassuk@uiowa.edu
Affiliations The University of Iowa, Iowa City, Iowa, United States of America, Department of Pediatrics, The University of Iowa, Iowa City, Iowa, United States of America, Interdisciplinary Program in Genetics, The University of Iowa, Iowa City, Iowa, United States of America, Department of Neurology, The University of Iowa, Iowa City, Iowa, United States of America, Roy J. and Lucille A. Carver College of Medicine, The University of Iowa, Iowa City, Iowa, United States of America, Interdisciplinary Graduate Program in Molecular and Cellular Biology, The University of Iowa, Iowa City, Iowa, United States of America
Competing Interests
The authors have declared that no competing interests exist.
Author Contributions
Conceived and designed the experiments: LP PG NU KT JRM AGB TM. Performed the experiments: LP SW JMS SNE HT AM KT TM LPS. Analyzed the data: LP TW DSR AAP JMS HCM GLC LPS KT JAW PJF HE BWD TS HT JKB TM. Contributed reagents/materials/analysis tools: AGB VBM PG KT TM JRM. Wrote the paper: LP VBM JRM AGB.