About the Authors

Marco A. S. Baptista

mbaptista@michaeljfox.org

Affiliation Research Programs, The Michael J. Fox Foundation for Parkinson’s Research, New York, New York, United States of America

Kuldip D. Dave

Affiliation Research Programs, The Michael J. Fox Foundation for Parkinson’s Research, New York, New York, United States of America

Mark A. Frasier

Affiliation Research Programs, The Michael J. Fox Foundation for Parkinson’s Research, New York, New York, United States of America

Todd B. Sherer

Affiliation Research Programs, The Michael J. Fox Foundation for Parkinson’s Research, New York, New York, United States of America

Melanie Greeley

Affiliation WIL Research, Ashland, Ohio, United States of America

Melissa J. Beck

Affiliation WIL Research, Ashland, Ohio, United States of America

Julie S. Varsho

Affiliation WIL Research, Ashland, Ohio, United States of America

George A. Parker

Affiliation WIL Research, Ashland, Ohio, United States of America

Cindy Moore

Affiliation Research Services, VA Medical Center/Portland and the Department of Behavioral Neuroscience and Pathology, Oregon Health and Science University, Portland, Oregon, United States of America

Madeline J. Churchill

Affiliation Research Services, VA Medical Center/Portland and the Department of Behavioral Neuroscience and Pathology, Oregon Health and Science University, Portland, Oregon, United States of America

Charles K. Meshul

Affiliation Research Services, VA Medical Center/Portland and the Department of Behavioral Neuroscience and Pathology, Oregon Health and Science University, Portland, Oregon, United States of America

Brian K. Fiske

Affiliation Research Programs, The Michael J. Fox Foundation for Parkinson’s Research, New York, New York, United States of America

Competing Interests

The authors MG, MJB, JSV, and GAP are from WIL Research, an entirely service organization. WIL Research has no financial or any other competing interests in the results of the studies reported in this manuscript and this does not alter our adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MASB KDD MAF TBS MG MJB JSV GAP CKM BKF. Performed the experiments: MG MJB JSV CM MJC CKM. Analyzed the data: MG MJB JSV MJC CKM. Contributed reagents/materials/analysis tools: GAP CKM. Wrote the manuscript: MASB MG MJB CKM.