About the Authors

Heather M. Bryan

heabry@gmail.com

Current address: Department of Geography, University of Victoria, Victoria, British Columbia, Canada

Affiliations Faculty of Veterinary Medicine, University of Calgary, Calgary, Alberta, Canada, Raincoast Conservation Foundation, Bella Bella, British Columbia, Canada

Chris T. Darimont

Affiliations Raincoast Conservation Foundation, Bella Bella, British Columbia, Canada, Department of Geography, University of Victoria, Victoria, British Columbia, Canada

Paul C. Paquet

Affiliations Raincoast Conservation Foundation, Bella Bella, British Columbia, Canada, Faculty of Environmental Design, University of Calgary, Calgary, Alberta, Canada

Katherine E. Wynne-Edwards

Affiliation Faculty of Veterinary Medicine, University of Calgary, Calgary, Alberta, Canada

Judit E. G. Smits

Affiliation Faculty of Veterinary Medicine, University of Calgary, Calgary, Alberta, Canada

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HMB CTD PCP JEGS. Performed the experiments: HMB CTD KEWE JEGS. Analyzed the data: HMB CTD JEGS KEWE. Contributed reagents/materials/analysis tools: CTD PCP KEWE JEGS. Wrote the paper: HMB CTD PCP KEWE JEGS.