About the Authors

James I. Kirkland

jameskirkland@utah.gov

Affiliation Utah Geological Survey, Salt Lake City, Utah, United States of America

Luis Alcalá

Affiliation Fundación Conjunto Paleontológico de Teruel-Dinópolis, Museo Aragonés de Paleontología, Teruel, Spain

Mark A. Loewen

Affiliations Natural History Museum of Utah, Salt Lake City, Utah, United States of America, Department of Geology & Geophysics, University of Utah, Salt Lake City, Utah, United States of America

Eduardo Espílez

Affiliation Fundación Conjunto Paleontológico de Teruel-Dinópolis, Museo Aragonés de Paleontología, Teruel, Spain

Luis Mampel

Affiliation Fundación Conjunto Paleontológico de Teruel-Dinópolis, Museo Aragonés de Paleontología, Teruel, Spain

Jelle P. Wiersma

Affiliations Natural History Museum of Utah, Salt Lake City, Utah, United States of America, Department of Geology & Geophysics, University of Utah, Salt Lake City, Utah, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JIK LA MAL. Performed the experiments: JIK MAL. Analyzed the data: JIK LA MAL. Wrote the paper: JIK LA MAL. Oversaw the entire Ariño project: LA. Co-directed the excavation: EE LM. Oversaw the preparation of all the fossil materials: EE. Assisted with character evaluations and in constructing many of the figures: JPW. Coordinated quarry mapping and photography: LM EE.