About the Authors

Alan M. Elder

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Davin M. Henderson

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Amy V. Nalls

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Jason M. Wilham

Affiliation Rocky Mountain Laboratories, National Institute for Allergy and Infectious Diseases, National Institutes of Health, Hamilton, Montana, United States of America

Byron W. Caughey

Affiliation Rocky Mountain Laboratories, National Institute for Allergy and Infectious Diseases, National Institutes of Health, Hamilton, Montana, United States of America

Edward A. Hoover

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Anthony E. Kincaid

Affiliation Medical Microbiology and Immunology, Creighton University, Omaha, Nebraska, United States of America

Jason C. Bartz

Affiliation Medical Microbiology and Immunology, Creighton University, Omaha, Nebraska, United States of America

Candace K. Mathiason

candace.mathiason@colostate.edu

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AME CKM JCB. Performed the experiments: AME AVN AEK. Analyzed the data: AME AEK CKM. Contributed reagents/materials/analysis tools: EAH DMH JCB AEK BWC JMW. Wrote the manuscript: AME CKM.