About the Authors

Kenneth Carpenter

Ken.Carpenter@ceu.edu

Affiliations Prehistoric Museum, Utah State University – Eastern, Price, Utah, United States of America, Geology Section, University of Colorado Museum, Boulder, Colorado, United States of America

Tony DiCroce

Affiliation Denver Museum of Nature and Science, Denver, Colorado, United States of America

Billy Kinneer

Affiliation Denver Museum of Nature and Science, Denver, Colorado, United States of America

Robert Simon

Affiliation Dinosaur Safaris Inc., Ashland, Virginia, United States of America

Competing Interests

Robert Simon is affiliated with the business Dinosaur Safaris Inc., Kenneth Carpenter gets a salary from Utah State University, and Billy Kinneer and Tony DiCroce have retirement pensions. There are no patents, products in development or marketed products to declare because this is a manuscript on vertebrate fossils. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: KC. Performed the experiments: TD BK. Contributed reagents/materials/analysis tools: RS. Wrote the paper: KC.