About the Authors

Larissa M. Williams

Affiliations Biology Department, Woods Hole Oceanographic Institution, Woods Hole, Massachusetts, United States of America, Biology Department, Bates College, Lewiston, Maine, United States of America

Alicia R. Timme-Laragy

Affiliation Biology Department, Woods Hole Oceanographic Institution, Woods Hole, Massachusetts, United States of America

Jared V. Goldstone

Affiliation Biology Department, Woods Hole Oceanographic Institution, Woods Hole, Massachusetts, United States of America

Andrew G. McArthur

Affiliation Andrew McArthur Consulting, Hamilton, Ontario, Canada

John J. Stegeman

Affiliation Biology Department, Woods Hole Oceanographic Institution, Woods Hole, Massachusetts, United States of America

Roxanna M. Smolowitz

Affiliation Department of Biology and Marine Biology, Roger Williams University, Bristol, Rhode Island, United States of America

Mark E. Hahn

mhahn@whoi.edu

Affiliation Biology Department, Woods Hole Oceanographic Institution, Woods Hole, Massachusetts, United States of America

Competing Interests

Dr. McArthur is the sole employee of Andrew McArthur Consulting. Dr. McArthur, a long-time colleague formerly at the Marine Biological Laboratory in Woods Hole, is a collaborator whose effort is paid for (by the grant to M.E.H.) as a consultant. The fact that Dr. McArthur collaborates with the authors as a private consultant does not alter their adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: LMW ARTL AGM JJS MEH. Performed the experiments: LMW ARTL JVG. Analyzed the data: LMW ARTL AGM JVG RMS. Contributed reagents/materials/analysis tools: JJS RMS MEH. Wrote the manuscript: LMW ARTL AGM JVG JJS RMS MEH.