About the Authors

Bethany J. Wilson

bethany.wilson@sydney.edu.au

Affiliation Faculty of Veterinary Science, The University of Sydney, Sydney, New South Wales, Australia

Frank W. Nicholas

Affiliation Faculty of Veterinary Science, The University of Sydney, Sydney, New South Wales, Australia

John W. James

Affiliation Faculty of Veterinary Science, The University of Sydney, Sydney, New South Wales, Australia

Claire M. Wade

Affiliation Faculty of Veterinary Science, The University of Sydney, Sydney, New South Wales, Australia

Herman W. Raadsma

Affiliation Faculty of Veterinary Science, The University of Sydney, Sydney, New South Wales, Australia

Peter C. Thomson

Affiliation Faculty of Veterinary Science, The University of Sydney, Sydney, New South Wales, Australia

Competing Interests

Please note that co-author Claire Wade is a PLOS ONE Editorial Board member. The authors declare that this does not alter their adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: BJW FWN JWJ CMW HWR PCT. Performed the experiments: BJW. Analyzed the data: BJW PCT JWJ FWN CMW. Contributed reagents/materials/analysis tools: PCT. Wrote the paper: BJW.