About the Authors

J. Berton C. Harris

johnbh@princeton.edu

Affiliations Fundación de Conservación Jocotoco, Quito, Ecuador, Woodrow Wilson School of Public and International Affairs, Princeton University, Princeton, New Jersey, United States of America

David G. Haskell

Affiliation Department of Biology, the University of the South, Sewanee, Tennessee, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JBCH DGH. Performed the experiments: JBCH. Analyzed the data: JBCH. Contributed reagents/materials/analysis tools: JBCH DGH. Wrote the manuscript: JBCH DGH.