About the Authors

Munenari Itoh

Contributed equally to this work with: Munenari Itoh, Noriko Umegaki-Arao

Current address: Department of Dermatology, the Jikei University School of Medicine, Tokyo, Japan

Affiliation Department of Dermatology, Columbia University, College of Physicians & Surgeons, New York, New York, United States of America

Noriko Umegaki-Arao

Contributed equally to this work with: Munenari Itoh, Noriko Umegaki-Arao

Affiliation Department of Dermatology, Columbia University, College of Physicians & Surgeons, New York, New York, United States of America

Zongyou Guo

Affiliation Department of Dermatology, Columbia University, College of Physicians & Surgeons, New York, New York, United States of America

Liang Liu

Affiliation Department of Dermatology, Columbia University, College of Physicians & Surgeons, New York, New York, United States of America

Claire A. Higgins

Affiliation Department of Dermatology, Columbia University, College of Physicians & Surgeons, New York, New York, United States of America

Angela M. Christiano

amc65@columbia.edu

Affiliations Department of Dermatology, Columbia University, College of Physicians & Surgeons, New York, New York, United States of America, Department of Genetics & Development, Columbia University, College of Physicians & Surgeons, New York, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MI AMC. Performed the experiments: MI NU ZG LL CAH. Analyzed the data: MI NU. Contributed reagents/materials/analysis tools: MI NU ZG. Wrote the manuscript: MI AMC.