About the Authors

David Rudel

rudeld@ecu.edu

Affiliation Department of Biology, East Carolina University, Greenville, North Carolina, United States of America

Chandler D. Douglas

Affiliation Department of Biology, East Carolina University, Greenville, North Carolina, United States of America

Ian M. Huffnagle

Affiliation Department of Biology, East Carolina University, Greenville, North Carolina, United States of America

John M. Besser

Affiliation Columbia Environmental Research Center, U.S. Geological Survey, Columbia, Missouri, United States of America

Christopher G. Ingersoll

Affiliation Columbia Environmental Research Center, U.S. Geological Survey, Columbia, Missouri, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DR CDD JMB CGI. Performed the experiments: CDD IMH DR. Analyzed the data: DR CDD JMB CGI. Contributed reagents/materials/analysis tools: DR JMB CGI. Wrote the manuscript: DR CDD IMH JMB CGI.