About the Authors

Yan Chen

Affiliations Pathogen Molecular Genetics Section, Laboratory of Human Bacterial Pathogenesis, National Institute of Allergy and Infectious Diseases, U.S. National Institutes of Health, Bethesda, Maryland, United States of America, Department of Infectious Diseases, Sir Run Run Shaw Hospital, College of Medicine, Zhejiang University, Hangzhou, China

Som S. Chatterjee

Affiliation Pathogen Molecular Genetics Section, Laboratory of Human Bacterial Pathogenesis, National Institute of Allergy and Infectious Diseases, U.S. National Institutes of Health, Bethesda, Maryland, United States of America

Stephen F. Porcella

Affiliation Genomics Unit, Research Technologies Branch, National Institute of Allergy and Infectious Diseases, U.S. National Institutes of Health, Hamilton, Montana, United States of America

Yun-Song Yu

Affiliation Department of Infectious Diseases, Sir Run Run Shaw Hospital, College of Medicine, Zhejiang University, Hangzhou, China

Michael Otto

motto@niaid.nih.gov

Affiliation Pathogen Molecular Genetics Section, Laboratory of Human Bacterial Pathogenesis, National Institute of Allergy and Infectious Diseases, U.S. National Institutes of Health, Bethesda, Maryland, United States of America

Competing Interests

Michael Otto is an Academic Editor for PLOS ONE. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SC SP YY MO. Performed the experiments: YC. Analyzed the data: SP MO. Wrote the manuscript: MO.