About the Authors

Sonia A. Kjos

Current address: Department of Biology, University of Minnesota, Duluth, Minnesota, United States of America

Affiliation Marshfield Clinic Research Foundation, Marshfield, Wisconsin, United States of America

Stephanie A. Irving

Current address: Center for Health Research, Kaiser Permanente, Portland, Oregon, United States of America

Affiliation Marshfield Clinic Research Foundation, Marshfield, Wisconsin, United States of America

Jennifer K. Meece

Affiliation Marshfield Clinic Research Foundation, Marshfield, Wisconsin, United States of America

Edward A. Belongia

belongia.edward@marshfieldclinic.org

Affiliation Marshfield Clinic Research Foundation, Marshfield, Wisconsin, United States of America

Competing Interests

EAB and JKM receive research funding from MedImmune, LLC. This does not alter the authors’ adherence to all PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: EAB SAK SAI. Performed the experiments: SAK SAI JKM. Analyzed the data: SAK SAI EAB. Contributed reagents/materials/analysis tools: JKM. Wrote the paper: SAK EAB.