About the Authors

Amy V. Nalls

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Erin McNulty

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Jenny Powers

Affiliation Biological Resource Management Division, National Park Service, Fort Collins, Colorado, United States of America

Davis M. Seelig

Affiliation Department of Veterinary Clinical Sciences, University of Minnesota, Saint Paul, Minnesota, United States of America

Clare Hoover

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Nicholas J. Haley

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Jeanette Hayes-Klug

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Kelly Anderson

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Paula Stewart

Affiliation The Roslin Institute and Royal (Dick) School of Veterinary Studies, University of Edinburgh, Easter Bush, Midlothian, Scotland, United Kingdom

Wilfred Goldmann

Affiliation The Roslin Institute and Royal (Dick) School of Veterinary Studies, University of Edinburgh, Easter Bush, Midlothian, Scotland, United Kingdom

Edward A. Hoover

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Candace K. Mathiason

Candace.Mathiason@colostate.edu

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AVN CKM WG. Performed the experiments: AVN CKM DMS PS. Analyzed the data: AVN CKM WG DMS. Contributed reagents/materials/analysis tools: EAH WG. Wrote the paper: CKM AVN. Performed necropsies: DMS NJH CEH. Animal care and management: JHK KA EM. Veterinary care and procedures: JP.