About the Authors

Brandon P. Hedrick

bhedrick@sas.upenn.edu

Affiliation Department of Earth and Environmental Science, School of Arts and Sciences, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Peter Dodson

Affiliations Department of Earth and Environmental Science, School of Arts and Sciences, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America, School of Veterinary Medicine, Department of Animal Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

Peter Dodson is a PLOS ONE editorial board member. This does not alter the authors' adherence to all of the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: BPH PD. Performed the experiments: BPH. Analyzed the data: BPH. Contributed reagents/materials/analysis tools: BPH PD. Wrote the paper: BPH PD.