About the Authors

Fergus G. Neville

fgn@st-and.ac.uk

Affiliation School of Medicine, University of St Andrews, St Andrews, Fife, United Kingdom

Damien J. Williams

Affiliation School of Medicine, University of St Andrews, St Andrews, Fife, United Kingdom

Christine A. Goodall

Affiliation Department of Oral Surgery, University of Glasgow Dental School, Glasgow, United Kingdom

Jeffrey S. Murer

Affiliation School of International Relations, University of St Andrews, St Andrews, Fife, United Kingdom

Peter D. Donnelly

Affiliation School of Medicine, University of St Andrews, St Andrews, Fife, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: FGN DJW CAG JSM PPD. Performed the experiments: FGN DJW CAG. Analyzed the data: FGN DJW CAG JSM PPD. Contributed reagents/materials/analysis tools: FGN DJW CAG JSM PPD. Wrote the paper: FGN DJW CAG JSM PPD.