About the Authors

Thomas A. Mendel

Affiliations Department of Ophthalmology, University of Virginia, Charlottesville, Virginia, United States of America, Department of Pathology, University of Virginia, Charlottesville, Virginia, United States of America

Erin B. D. Clabough

Current address: Department of Biology, Randolph-Macon College, Ashland, Virginia, United States of America

Affiliation Department of Ophthalmology, University of Virginia, Charlottesville, Virginia, United States of America

David S. Kao

Affiliation Department of Ophthalmology, University of Virginia, Charlottesville, Virginia, United States of America

Tatiana N. Demidova-Rice

Current address: E.L. Steele Laboratory for Tumor Biology, Massachusetts General Hospital, Harvard Medical School, Boston, Massachusetts, United States of America

Affiliation Department of Molecular Physiology and Pharmacology, Tufts University, Boston, Massachusetts, United States of America

Jennifer T. Durham

Affiliation Department of Molecular Physiology and Pharmacology, Tufts University, Boston, Massachusetts, United States of America

Brendan C. Zotter

Affiliation Department of Ophthalmology, University of Virginia, Charlottesville, Virginia, United States of America

Scott A. Seaman

Affiliation Department of Biomedical Engineering, University of Virginia, Charlottesville, Virginia, United States of America

Stephen M. Cronk

Affiliation Department of Biomedical Engineering, University of Virginia, Charlottesville, Virginia, United States of America

Elizabeth P. Rakoczy

Affiliation Center for Ophthalmology and Visual Sciences, Department of Molecular Ophthalmology, The University of Western Australia, Nedlands, Western Australia, Australia

Adam J. Katz

Current address: Department of Plastic Surgery, University of Florida, Gainesville, Florida, United States of America

Affiliation Department of Plastic Surgery, University of Virginia, Charlottesville, Virginia, United States of America

Ira M. Herman

Affiliation Department of Molecular Physiology and Pharmacology, Tufts University, Boston, Massachusetts, United States of America

Shayn M. Peirce

Affiliation Department of Biomedical Engineering, University of Virginia, Charlottesville, Virginia, United States of America

Paul A. Yates

pay2x@virginia.edu

Affiliation Department of Ophthalmology, University of Virginia, Charlottesville, Virginia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TAM EBDC TND JTD BCZ SAS SMC IMH SMP PAY. Performed the experiments: TAM EBDC DSK TND JTD BCZ SAS SMC. Analyzed the data: TAM EBDC TND JTD BCZ SAS SMC IMH SMP PAY. Contributed reagents/materials/analysis tools: EPR AJK IMH. Wrote the paper: TAM EBDC TND JTD IMH SMP PAY.