About the Authors

David A. Davis

Contributed equally to this work with: David A. Davis, Marco Bortolato, Caleb E. Finch, Todd E. Morgan

Affiliation Davis School of Gerontology, USC, Los Angeles, California, United States of America

Marco Bortolato

Contributed equally to this work with: David A. Davis, Marco Bortolato, Caleb E. Finch, Todd E. Morgan

Affiliation Dept. of Pharmacology and Toxicology, School of Pharmacy, University of Kansas, Lawrence, Kansas, United States of America

Sean C. Godar

Affiliation School of Pharmacy, USC, Los Angeles, California, United States of America

Thomas K. Sander

Affiliation Dornsife College of Letters, Arts and Sciences, USC, Los Angeles, California, United States of America

Nahoko Iwata

Affiliation Davis School of Gerontology, USC, Los Angeles, California, United States of America

Payam Pakbin

Affiliation Viterbi School of Engineering, USC, Los Angeles, California, United States of America

Jean C. Shih

Affiliation School of Pharmacy, USC, Los Angeles, California, United States of America

Kiros Berhane

Affiliation Keck School of Medicine, USC, Los Angeles, California, United States of America

Rob McConnell

Affiliation Keck School of Medicine, USC, Los Angeles, California, United States of America

Constantinos Sioutas

Affiliation Viterbi School of Engineering, USC, Los Angeles, California, United States of America

Caleb E. Finch

Contributed equally to this work with: David A. Davis, Marco Bortolato, Caleb E. Finch, Todd E. Morgan

cefinch@usc.edu

Affiliations Davis School of Gerontology, USC, Los Angeles, California, United States of America, Dept. of Neurobiology, Dornsife College, USC, Los Angeles, California, United States of America

Todd E. Morgan

Contributed equally to this work with: David A. Davis, Marco Bortolato, Caleb E. Finch, Todd E. Morgan

Affiliation Davis School of Gerontology, USC, Los Angeles, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TEM CEF. Performed the experiments: DAD MB SCG TKS NI PP. Analyzed the data: RM KB MB. Contributed reagents/materials/analysis tools: CS. Wrote the paper: DAD MC RM CEF TEM. Behavior: MB SCG JCS.