About the Authors

Sergey Lyskov

Contributed equally to this work with: Sergey Lyskov, Fang-Chieh Chou

Affiliation Department of Chemical and Biomolecular Engineering, The Johns Hopkins University, Baltimore, Maryland, United States of America

Fang-Chieh Chou

Contributed equally to this work with: Sergey Lyskov, Fang-Chieh Chou

Affiliation Department of Biochemistry, Stanford University School of Medicine, Stanford, California, United States of America

Shane Ó. Conchúir

Affiliations California Institute for Quantitative Biomedical Research, University of California San Francisco, San Francisco, California, United States of America, Department of Bioengineering and Therapeutic Sciences, University of California San Francisco, San Francisco, California, United States of America

Bryan S. Der

Affiliation Department of Biochemistry and Biophysics, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Kevin Drew

Affiliation Department of Biology, Center for Genomics and Systems Biology, New York University, New York, New York, United States of America

Daisuke Kuroda

Affiliation Department of Chemical and Biomolecular Engineering, The Johns Hopkins University, Baltimore, Maryland, United States of America

Jianqing Xu

Affiliation Department of Chemical and Biomolecular Engineering, The Johns Hopkins University, Baltimore, Maryland, United States of America

Brian D. Weitzner

Affiliation Department of Chemical and Biomolecular Engineering, The Johns Hopkins University, Baltimore, Maryland, United States of America

P. Douglas Renfrew

Affiliation Department of Biology, Center for Genomics and Systems Biology, New York University, New York, New York, United States of America

Parin Sripakdeevong

Affiliation Biophysics Program, Stanford University, Stanford, California, United States of America

Benjamin Borgo

Affiliation Department of Genetics, Washington University in St. Louis, St. Louis, Missouri, United States of America

James J. Havranek

Affiliation Department of Genetics, Washington University in St. Louis, St. Louis, Missouri, United States of America

Brian Kuhlman

Affiliation Department of Biochemistry and Biophysics, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Tanja Kortemme

Affiliations California Institute for Quantitative Biomedical Research, University of California San Francisco, San Francisco, California, United States of America, Department of Bioengineering and Therapeutic Sciences, University of California San Francisco, San Francisco, California, United States of America, Graduate Group in Biophysics, University of California San Francisco, San Francisco, California, United States of America

Richard Bonneau

Affiliations Department of Biology, Center for Genomics and Systems Biology, New York University, New York, New York, United States of America, Computer Science Department, Courant Institute of Mathematical Sciences, New York University, New York, New York, United States of America

Jeffrey J. Gray

jgray@jhu.edu (JG); rhiju@stanford.edu (RD)

Affiliations Department of Chemical and Biomolecular Engineering, The Johns Hopkins University, Baltimore, Maryland, United States of America, Program in Molecular Biophysics, The Johns Hopkins University, Baltimore, Maryland, United States of America

Rhiju Das

jgray@jhu.edu (JG); rhiju@stanford.edu (RD)

Affiliations Department of Biochemistry, Stanford University School of Medicine, Stanford, California, United States of America, Department of Physics, Stanford University, Stanford, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Contributed reagents/materials/analysis tools: SL FC SÓC BD KD DK JX BW PDR PS BB JH BK TK RB JG RD. Wrote the paper: SL FC SÓC BD KD DK JX BW PDR PS BB JH BK TK RB JG RD.