About the Authors

Hiromi Ikeda

Affiliation Section on Model Synaptic Systems, Laboratory of Molecular Physiology, National Institute on Alcohol Abuse and Alcoholism, National Institutes of Health, Rockville, Maryland, United States of America

Alison H. Delargy

Affiliation Section on Model Synaptic Systems, Laboratory of Molecular Physiology, National Institute on Alcohol Abuse and Alcoholism, National Institutes of Health, Rockville, Maryland, United States of America

Tohei Yokogawa

Affiliation Unit on Behavioral Neurogenetics, National Institute of Child Health and Development, National Institutes of Health, Bethesda, Maryland, United States of America

Jason M. Urban

Affiliation Section on Model Synaptic Systems, Laboratory of Molecular Physiology, National Institute on Alcohol Abuse and Alcoholism, National Institutes of Health, Rockville, Maryland, United States of America

Harold A. Burgess

Affiliation Unit on Behavioral Neurogenetics, National Institute of Child Health and Development, National Institutes of Health, Bethesda, Maryland, United States of America

Fumihito Ono

onof@mail.nih.gov

Affiliation Section on Model Synaptic Systems, Laboratory of Molecular Physiology, National Institute on Alcohol Abuse and Alcoholism, National Institutes of Health, Rockville, Maryland, United States of America

Competing Interests

One of the co-authors, Harold Burgess, is a PLOS ONE Editorial Board member. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: HI FO. Performed the experiments: HI AHD. Analyzed the data: HI HAB. Contributed reagents/materials/analysis tools: AHD TY JMU HAB. Wrote the paper: HI FO.