About the Authors

Fang K. Du

dufang325@gmail.com (FKD); RWU@hes.hmc.psu.edu (RW)

Affiliation Center for Computational Biology, National Engineering Laboratory for Tree Breeding, College of Biological Sciences and Biotechnology, Beijing Forestry University, Beijing, China

Fang Xu

Affiliation Center for Computational Biology, National Engineering Laboratory for Tree Breeding, College of Biological Sciences and Biotechnology, Beijing Forestry University, Beijing, China

Hong Qu

Affiliation Center for Bioinformatics, National Laboratory of Protein Engineering and Plant Genetic Engineering, College of Life Sciences, Peking University, Beijing, China

Sisi Feng

Affiliation Center for Computational Biology, National Engineering Laboratory for Tree Breeding, College of Biological Sciences and Biotechnology, Beijing Forestry University, Beijing, China

Jijun Tang

Affiliation Department of Computer Science and Engineering, University of South Carolina, Columbia, South Carolina, United States of America

Rongling Wu

dufang325@gmail.com (FKD); RWU@hes.hmc.psu.edu (RW)

Affiliation Center for Computational Biology, National Engineering Laboratory for Tree Breeding, College of Biological Sciences and Biotechnology, Beijing Forestry University, Beijing, China

Competing Interests

Co-author Rongling Wu is a PLOS ONE Editorial Board member; this does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: FKD RW. Performed the experiments: FKD FX SF. Analyzed the data: FKD. Contributed reagents/materials/analysis tools: HQ JT. Wrote the paper: FKD.