About the Authors

Leonie R. Clitherow

Current address: School of Geography, Earth and Environmental Sciences, University of Birmingham, Edgbaston, Birmingham, United Kingdom

Affiliation Faculty of Biological Science/water@leeds, University of Leeds, Leeds, United Kingdom

Jonathan L. Carrivick

Affiliation School of Geography/water@leeds, University of Leeds, Leeds, United Kingdom

Lee E. Brown

l.brown@leeds.ac.uk

Affiliation School of Geography/water@leeds, University of Leeds, Leeds, United Kingdom

Competing Interests

LRC was funded by the Santander Full Fee Masters Scholarship. There are no patents, products in development or marketed products to declare. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: LEB LRC JLC. Performed the experiments: LRC. Analyzed the data: LRC LEB. Contributed reagents/materials/analysis tools: LRC LEB JLC. Wrote the paper: LRC LEB.