About the Authors

Akinori Mitani

Affiliations Department of Neurophysiology, Graduate School of Medicine, Juntendo University, Bunkyo, Tokyo, Japan, Faculty of Medicine, The University of Tokyo, Bunkyo, Tokyo, Japan, Neuroscience Graduate Program, University of California San Diego, La Jolla, California, United States of America

Ryo Sasaki

Affiliations Department of Neurophysiology, Graduate School of Medicine, Juntendo University, Bunkyo, Tokyo, Japan, Department of Brain and Cognitive Sciences, Center for Visual Science, University of Rochester, Rochester, New York, United States of America

Masafumi Oizumi

Affiliations Laboratory for Mathematical Neuroscience, RIKEN Brain Science Institute, Wako, Saitama, Japan, Department of Psychiatry, University of Wisconsin, Madison, Wisconsin, United States of America

Takanori Uka

uka@juntendo.ac.jp

Affiliation Department of Neurophysiology, Graduate School of Medicine, Juntendo University, Bunkyo, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AM RS MO TU. Performed the experiments: AM RS. Analyzed the data: AM RS. Wrote the paper: AM RS MO TU.