About the Authors

Yasue Horiuchi

Contributed equally to this work with: Yasue Horiuchi, Maya Ishikawa

Affiliation Department of Medical Genetics, Majors of Medical Sciences, Graduate School of Comprehensive Human Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan

Maya Ishikawa

Contributed equally to this work with: Yasue Horiuchi, Maya Ishikawa

Affiliation Department of Medical Genetics, Majors of Medical Sciences, Graduate School of Comprehensive Human Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan

Nobuko Kaito

Affiliation Department of Medical Genetics, Majors of Medical Sciences, Graduate School of Comprehensive Human Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan

Yoshimi Iijima

Affiliation Department of Medical Genetics, Majors of Medical Sciences, Graduate School of Comprehensive Human Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan

Yoshiko Tanabe

Affiliation Department of Medical Genetics, Majors of Medical Sciences, Graduate School of Comprehensive Human Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan

Hiroki Ishiguro

Affiliation Department of Medical Genetics, Majors of Medical Sciences, Graduate School of Comprehensive Human Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan

Tadao Arinami

tarinami@md.tsukuba.ac.jp

Affiliation Department of Medical Genetics, Majors of Medical Sciences, Graduate School of Comprehensive Human Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YH HI TA. Performed the experiments: YH MI NK YT. Analyzed the data: YH MI YI. Contributed reagents/materials/analysis tools: HI. Wrote the paper: YH TA.