About the Authors

Samuel Herberg

Affiliations Charlie Norwood VA Medical Center, Augusta, Georgia, United States of America, Department of Cellular Biology and Anatomy, Georgia Regents University, Augusta, Georgia, United States of America

Xingming Shi

Affiliations Department of Pathology, Georgia Regents University, Augusta, Georgia, United States of America, Institute of Molecular Medicine and Genetics, Georgia Regents University, Augusta, Georgia, United States of America, Institute of Regenerative and Reparative Medicine, Georgia Regents University, Augusta, Georgia, United States of America

Maribeth H. Johnson

Affiliation Department of Biostatistics and Epidemiology, Georgia Regents University, Augusta, Georgia, United States of America

Mark W. Hamrick

Affiliations Department of Cellular Biology and Anatomy, Georgia Regents University, Augusta, Georgia, United States of America, Department of Orthopaedic Surgery, Georgia Regents University, Augusta, Georgia, United States of America, Institute of Molecular Medicine and Genetics, Georgia Regents University, Augusta, Georgia, United States of America, Institute of Regenerative and Reparative Medicine, Georgia Regents University, Augusta, Georgia, United States of America

Carlos M. Isales

Affiliations Department of Cellular Biology and Anatomy, Georgia Regents University, Augusta, Georgia, United States of America, Department of Orthopaedic Surgery, Georgia Regents University, Augusta, Georgia, United States of America, Institute of Molecular Medicine and Genetics, Georgia Regents University, Augusta, Georgia, United States of America, Institute of Regenerative and Reparative Medicine, Georgia Regents University, Augusta, Georgia, United States of America

William D. Hill

whill@gru.edu

Affiliations Charlie Norwood VA Medical Center, Augusta, Georgia, United States of America, Department of Cellular Biology and Anatomy, Georgia Regents University, Augusta, Georgia, United States of America, Department of Orthopaedic Surgery, Georgia Regents University, Augusta, Georgia, United States of America, Institute of Molecular Medicine and Genetics, Georgia Regents University, Augusta, Georgia, United States of America, Institute of Regenerative and Reparative Medicine, Georgia Regents University, Augusta, Georgia, United States of America

Competing Interests

The authors would like to declare that their co-author Dr. Xingming Shi is a PLOS ONE Editorial Board member. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SH XS MHJ MWH CMI WDH. Performed the experiments: SH. Analyzed the data: SH MHJ WDH. Contributed reagents/materials/analysis tools: XS. Wrote the paper: SH XS WDH.