About the Authors

Jean A. Hall

Jean.Hall@oregonstate.edu

Affiliation Department of Biomedical Sciences, College of Veterinary Medicine, Oregon State University, Corvallis, Oregon, United States of America

Gerd Bobe

Affiliations Department of Animal and Rangeland Sciences, College of Agricultural Sciences, Oregon State University, Corvallis, Oregon, United States of America, Linus Pauling Institute, Oregon State University, Corvallis, Oregon, United States of America

Janice K. Hunter

Current address: Deep Springs College, Dyer, Nevada, United States of America

Affiliation Department of Animal and Rangeland Sciences, College of Agricultural Sciences, Oregon State University, Corvallis, Oregon, United States of America

William R. Vorachek

Affiliation Department of Biomedical Sciences, College of Veterinary Medicine, Oregon State University, Corvallis, Oregon, United States of America

Whitney C. Stewart

Current address: Texas AgriLife Research, San Angelo, Texas, United States of America

Affiliation Department of Animal and Rangeland Sciences, College of Agricultural Sciences, Oregon State University, Corvallis, Oregon, United States of America

Jorge A. Vanegas

Affiliation Department of Clinical Sciences, College of Veterinary Medicine, Oregon State University, Corvallis, Oregon, United States of America

Charles T. Estill

Affiliations Department of Animal and Rangeland Sciences, College of Agricultural Sciences, Oregon State University, Corvallis, Oregon, United States of America, Department of Clinical Sciences, College of Veterinary Medicine, Oregon State University, Corvallis, Oregon, United States of America

Wayne D. Mosher

Affiliation Department of Animal and Rangeland Sciences, College of Agricultural Sciences, Oregon State University, Corvallis, Oregon, United States of America

Gene J. Pirelli

Affiliation Department of Animal and Rangeland Sciences, College of Agricultural Sciences, Oregon State University, Corvallis, Oregon, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JAH WDM GJP. Performed the experiments: JAH JKH WRV WCS JAV CTE WDM GJP. Analyzed the data: JAH GB. Contributed reagents/materials/analysis tools: JAH GB. Wrote the paper: JAH GB CTE GJP.