About the Authors

Anupama Tiwari

Affiliation Department of Animal Science, Center for Reproductive Biology and Health, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Jill A. Hadley

Affiliation Department of Animal Science, Center for Reproductive Biology and Health, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Gilbert L. Hendricks III

Affiliation Department of Animal Science, Center for Reproductive Biology and Health, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Robert G. Elkin

Affiliation Department of Animal Science, Center for Reproductive Biology and Health, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Timothy Cooper

Affiliation Department of Comparative Medicine, Penn State College of Medicine, Penn State Hershey Medical Center, Hershey, Pennsylvania, United States of America

Ramesh Ramachandran

RameshR@psu.edu

Affiliation Department of Animal Science, Center for Reproductive Biology and Health, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AT JAH RR. Performed the experiments: AT JAH GLH RGE TKC RR. Analyzed the data: AT JAH TKC RR. Wrote the paper: AT RR.