About the Authors

Jennifer M. Loftis

loftisj@ohsu.edu

Affiliations Research and Development, Portland VA Medical Center, Portland, Oregon, United States of America, Department of Psychiatry, School of Medicine, Oregon Health and Science University, Portland, Oregon, United States of America

Clare J. Wilhelm

Affiliations Research and Development, Portland VA Medical Center, Portland, Oregon, United States of America, Department of Psychiatry, School of Medicine, Oregon Health and Science University, Portland, Oregon, United States of America

Arthur A. Vandenbark

Affiliations Research and Development, Portland VA Medical Center, Portland, Oregon, United States of America, Department of Neurology, School of Medicine, Oregon Health and Science University, Portland, Oregon, United States of America, Department of Molecular Microbiology and Immunology, School of Medicine, Oregon Health and Science University, Portland, Oregon, United States of America

Marilyn Huckans

Affiliations Research and Development, Portland VA Medical Center, Portland, Oregon, United States of America, Behavioral Health & Clinical Neurosciences Division, Portland VA Medical Center, Portland, Oregon, United States of America, Department of Psychiatry, School of Medicine, Oregon Health and Science University, Portland, Oregon, United States of America

Competing Interests

The authors have read the journal's policy and have the following conflicts: the Department of Veterans Affairs and Oregon Health & Science University (OHSU) own a technology used in this research study. AAV has stock options in Virogenomics/Artielle, a company that has licensed the technology and may have an interest in the results of this study. The Department of Veterans Affairs, OHSU, and JML, AAV, and MH have rights to royalties from the licensing agreement with Artielle. These potential conflicts of interest have been reviewed and managed by the Conflict of Interest Committees at the Portland VA Medical Center and OHSU. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Provided critical review of the manuscript: CJW AAV MH. Conceived and designed the experiments: JML AAV MH. Performed the experiments: JML CJW. Analyzed the data: JML CJW MH. Contributed reagents/materials/analysis tools: AAV JML. Wrote the paper: JML.