About the Authors

Kai Wu

kai.wu.22@gmail.com

Affiliations Department of Nuclear Medicine and Radiology, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Japan, Department of Biomedical Engineering, School of Materials Science and Engineering, South China University of Technology, Guangzhou, China

Yasuyuki Taki

Affiliations Department of Nuclear Medicine and Radiology, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Japan, Division of Developmental Cognitive Neuroscience, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Japan

Kazunori Sato

Affiliation Department of Nuclear Medicine and Radiology, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Japan

Hiroshi Hashizume

Affiliation Division of Developmental Cognitive Neuroscience, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Japan

Yuko Sassa

Affiliation Division of Developmental Cognitive Neuroscience, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Japan

Hikaru Takeuchi

Affiliation Smart Ageing International Research Centre, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Japan

Benjamin Thyreau

Affiliation Division of Developmental Cognitive Neuroscience, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Japan

Yong He

Affiliation State Key Laboratory of Cognitive Neuroscience and Learning, Beijing Normal University, Beijing, China

Alan C. Evans

Affiliation McConnell Brain Imaging Centre, Montreal Neurological Institute, McGill University, Montreal, Quebec, Canada

Xiaobo Li

Affiliation Department of Radiology, Albert Einstein College of Medicine, Yeshiva University, Bronx, New York, United States of America

Ryuta Kawashima

Affiliations Division of Developmental Cognitive Neuroscience, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Japan, Smart Ageing International Research Centre, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Japan, Department of Functional Brain Imaging, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Japan

Hiroshi Fukuda

Affiliation Department of Nuclear Medicine and Radiology, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Japan

Competing Interests

One of the co-authors (Dr. Yong He) currently serves as an editor for PLOS ONE. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Revised the manuscript: YH ACE XL RK HF. Contributed to data collection and analysis: YS HH HT BT. Conceived and designed the experiments: KW YT KS HH YS HT BT. Performed the experiments: KW. Analyzed the data: KW. Contributed reagents/materials/analysis tools: KW YT KS. Wrote the paper: KW.