About the Authors

Qing Sang

Contributed equally to this work with: Qing Sang, Xukun Yan, Huan Wang

Affiliations State Key Laboratory of Genetic Engineering and MOE Key Laboratory of Contemporary Anthropology, School of Life Sciences, Fudan University, Shanghai, People's Republic of China, Institutes of Biomedical Sciences, Fudan University, Shanghai, People's Republic of China

Xukun Yan

Contributed equally to this work with: Qing Sang, Xukun Yan, Huan Wang

Affiliations Department of Otolaryngology, Eye & ENT Hospital, Fudan University, Shanghai, People's Republic of China, Department of Otolaryngology, Chinese PLA General Hospital, Beijing, People's Republic of China

Huan Wang

Contributed equally to this work with: Qing Sang, Xukun Yan, Huan Wang

Affiliations State Key Laboratory of Genetic Engineering and MOE Key Laboratory of Contemporary Anthropology, School of Life Sciences, Fudan University, Shanghai, People's Republic of China, Institutes of Biomedical Sciences, Fudan University, Shanghai, People's Republic of China

Ruizhi Feng

Affiliations State Key Laboratory of Genetic Engineering and MOE Key Laboratory of Contemporary Anthropology, School of Life Sciences, Fudan University, Shanghai, People's Republic of China, Institutes of Biomedical Sciences, Fudan University, Shanghai, People's Republic of China

Xiang Fei

Affiliation Fudan-VARI Center for Genetic Epidemiology, Fudan University, Shanghai, People's Republic of China

Duan Ma

Affiliation Department of Otolaryngology, Eye & ENT Hospital, Fudan University, Shanghai, People's Republic of China

Qinghe Xing

Affiliations Institutes of Biomedical Sciences, Fudan University, Shanghai, People's Republic of China, Bio-X Center, Key Laboratory for the Genetics of Developmental and Neuropsychiatric Disorders, Ministry of Education, Shanghai Jiao Tong University, Shanghai, People's Republic of China

Qiaoli Li

Affiliations Institutes of Biomedical Sciences, Fudan University, Shanghai, People's Republic of China, Bio-X Center, Key Laboratory for the Genetics of Developmental and Neuropsychiatric Disorders, Ministry of Education, Shanghai Jiao Tong University, Shanghai, People's Republic of China

Xinzhi Zhao

Affiliations Institutes of Biomedical Sciences, Fudan University, Shanghai, People's Republic of China, Bio-X Center, Key Laboratory for the Genetics of Developmental and Neuropsychiatric Disorders, Ministry of Education, Shanghai Jiao Tong University, Shanghai, People's Republic of China

Li Jin

Affiliation State Key Laboratory of Genetic Engineering and MOE Key Laboratory of Contemporary Anthropology, School of Life Sciences, Fudan University, Shanghai, People's Republic of China

Lin He

Affiliations Institutes of Biomedical Sciences, Fudan University, Shanghai, People's Republic of China, Bio-X Center, Key Laboratory for the Genetics of Developmental and Neuropsychiatric Disorders, Ministry of Education, Shanghai Jiao Tong University, Shanghai, People's Republic of China

Huawei Li

hwli@shmu.edu.cn (HWL); wangleiwanglei@fudan.edu.cn. (LW)

Affiliation Department of Otolaryngology, Eye & ENT Hospital, Fudan University, Shanghai, People's Republic of China

Lei Wang

hwli@shmu.edu.cn (HWL); wangleiwanglei@fudan.edu.cn. (LW)

Affiliations State Key Laboratory of Genetic Engineering and MOE Key Laboratory of Contemporary Anthropology, School of Life Sciences, Fudan University, Shanghai, People's Republic of China, Institutes of Biomedical Sciences, Fudan University, Shanghai, People's Republic of China, Bio-X Center, Key Laboratory for the Genetics of Developmental and Neuropsychiatric Disorders, Ministry of Education, Shanghai Jiao Tong University, Shanghai, People's Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Edited the manuscript: LW QS XQH LH HWL DM. Conceived and designed the experiments: LW QS HWL XKY. Performed the experiments: QS HW XKY. Analyzed the data: QS HW XKY RZF XF LW. Contributed reagents/materials/analysis tools: DM QHX LH QLL XZZ LJ. Wrote the paper: QS LW.