About the Authors

Tomokazu Fukuda

Contributed equally to this work with: Tomokazu Fukuda, Yasushi Kino

Affiliation Graduate School of Agricultural Sciences, Tohoku University, Sendai, Miyagi, Japan

Yasushi Kino

Contributed equally to this work with: Tomokazu Fukuda, Yasushi Kino

Affiliation Graduate School of Science, Tohoku University, Sendai, Miyagi, Japan

Yasuyuki Abe

Affiliation Graduate School of Science and Engineering, Yamagata University, Tsuruoka, Yamagata, Japan

Hideaki Yamashiro

Affiliation Faculty of Agriculture, Niigata University, Niigata, Niigata, Japan

Yoshikazu Kuwahara

Affiliation Department of Pathology, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Miyagi, Japan

Hidekazu Nihei

Affiliation Graduate School of Science, Tohoku University, Sendai, Miyagi, Japan

Yosuke Sano

Affiliation Graduate School of Science, Tohoku University, Sendai, Miyagi, Japan

Ayumi Irisawa

Affiliation Graduate School of Science, Tohoku University, Sendai, Miyagi, Japan

Tsutomu Shimura

Affiliation Department of Pathology, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Miyagi, Japan

Motoi Fukumoto

Affiliation Department of Pathology, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Miyagi, Japan

Hisashi Shinoda

Affiliation Graduate School of Dentistry, Tohoku University, Sendai, Miyagi, Japan

Yuichi Obata

Affiliation RIKEN BioResource Center (BRC), Tsukuba, Ibaraki, Japan

Shin Saigusa

Affiliation National Institute of Radiological Sciences, Chiba, Chiba, Japan

Tsutomu Sekine

Affiliation Center for the Advancement of Higher Education, Tohoku University, Sendai, Miyagi, Japan

Emiko Isogai

Affiliation Graduate School of Agricultural Sciences, Tohoku University, Sendai, Miyagi, Japan

Manabu Fukumoto

fukumoto@idac.tohoku.ac.jp

Affiliation Department of Pathology, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Miyagi, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: Manabu Fukumoto. Performed the experiments: TF Y. Kino YA HY HN YS AI T. Sekine EI Manabu Fukumoto. Analyzed the data: Y. Kino T. Sekine SS. Contributed reagents/materials/analysis tools: Y. Kuwahara T. Shimura Motoi Fukumoto HS YO M. Fukumoto. Wrote the paper: TF Y. Kino T. Sekine Manabu Fukumoto.