About the Authors

Ruth L. Angell

Affiliation Faculty of Biological Sciences, University of Leeds, Leeds, West Yorkshire, United Kingdom

Roger K. Butlin

Affiliation Dept. Animal and Plant Sciences, University of Sheffield, Sheffield, South Yorkshire, United Kingdom

John D. Altringham

j.d.altringham@leeds.ac.uk

Affiliation Faculty of Biological Sciences, University of Leeds, Leeds, West Yorkshire, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JDA RKB RLA. Performed the experiments: RLA JDA. Analyzed the data: RLA RKB. Wrote the paper: RLA JDA RKB.