About the Authors

Xueming Tang

Affiliations Biotechnology Research Institute, Shanghai Academy of Agricultural Sciences, Shanghai, People’s Republic of China, Key Laboratory of Agricultural Genetics and Breeding, Shanghai Academy of Agricultural Sciences, Shanghai, People’s Republic of China

Fangting Han

Affiliation College of Life and Environment Sciences, Shanghai Normal University, Shanghai, People’s Republic of China

Kai Zhao

kzhao118@163.com

Affiliations Biotechnology Research Institute, Shanghai Academy of Agricultural Sciences, Shanghai, People’s Republic of China, Key Laboratory of Agricultural Genetics and Breeding, Shanghai Academy of Agricultural Sciences, Shanghai, People’s Republic of China

Yan Xu

Affiliation College of Life and Environment Sciences, Shanghai Normal University, Shanghai, People’s Republic of China

Xiao Wu

Affiliations Biotechnology Research Institute, Shanghai Academy of Agricultural Sciences, Shanghai, People’s Republic of China, Key Laboratory of Agricultural Genetics and Breeding, Shanghai Academy of Agricultural Sciences, Shanghai, People’s Republic of China

Jinbin Wang

Affiliations Biotechnology Research Institute, Shanghai Academy of Agricultural Sciences, Shanghai, People’s Republic of China, Key Laboratory of Agricultural Genetics and Breeding, Shanghai Academy of Agricultural Sciences, Shanghai, People’s Republic of China

Lingxi Jiang

Affiliations Biotechnology Research Institute, Shanghai Academy of Agricultural Sciences, Shanghai, People’s Republic of China, Key Laboratory of Agricultural Genetics and Breeding, Shanghai Academy of Agricultural Sciences, Shanghai, People’s Republic of China

Wei Shi

Affiliation College of Life and Environment Sciences, Shanghai Normal University, Shanghai, People’s Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: XMT FTH KZ. Performed the experiments: XMT KZ FTH. Analyzed the data: XMT FTH KZ. Contributed reagents/materials/analysis tools: YX XW JBW LXJ WS. Wrote the paper: XMT FTH. Development of modeling approach: XW KZ. Extensive editorial comments, insight into results: XW KZ.