About the Authors

Masato Hosokawa

Affiliation Department of Dementia and Higher Brain Function, Tokyo Metropolitan Institute of Medical Science, Tokyo, Japan

Tetsuaki Arai

Affiliations Department of Dementia and Higher Brain Function, Tokyo Metropolitan Institute of Medical Science, Tokyo, Japan, Department of Psychiatry, Graduate School of Comprehensive Human Sciences, University of Tsukuba, Tsukuba, Japan

Masami Masuda-Suzukake

Affiliation Department of Pathology and Cell Biology, Tokyo Metropolitan Institute of Medical Science, Tokyo, Japan

Takashi Nonaka

Affiliation Department of Pathology and Cell Biology, Tokyo Metropolitan Institute of Medical Science, Tokyo, Japan

Makiko Yamashita

Affiliation Department of Pathology and Cell Biology, Tokyo Metropolitan Institute of Medical Science, Tokyo, Japan

Haruhiko Akiyama

Affiliation Department of Dementia and Higher Brain Function, Tokyo Metropolitan Institute of Medical Science, Tokyo, Japan

Masato Hasegawa

Affiliation Department of Pathology and Cell Biology, Tokyo Metropolitan Institute of Medical Science, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: M. Hosokawa TA HA. Performed the experiments: M. Hosokawa MM-S. Analyzed the data: M. Hosokawa MM-S TN MY HA M. Hasegawa. Contributed reagents/materials/analysis tools: MM-S TN M. Hasegawa. Wrote the paper: M. Hosokawa TA HA.